HYDROLOGIC AND HYDRAULIC DRIVERS OF THE UPPER MISSISSIPPI RIVER ECOSYSTEM Charles Theiling U.S. Army Corps of Engineers, Rock Island District, PO Box 2004, Clock Tower Bldg., Rock Island, IL 61204 River hydrology, in the broadest sense, is perhaps the strongest driver of floodplain river ecosystems. Hydrologic and hydraulic (H&H) processes drive sediment transport which shapes the channels and backwaters of aquatic habitats and forms the terrestrial geomorphic template that supports diverse plant and animal communities. River flow also transports dissolved nutrients and organic matter, including large woody debris, coarse and fine particulates, gametes, eggs, larvae, and invertebrates. H&H processes are important factors determining plant and animal habitats, especially soil moisture gradients ranging from permanent aquatic to rarely flooded and that support a range of aquatic to terrestrial upland plant species in relatively distinct community types along inundation gradients. Rheophylic fishes and macroinvertebrates exploit swift channels, while lentic fishes use off- channel aquatic habitat, and diving/swimming birds eat plants, invertebrates, or fishes in deepwater while dabblers and waders eat plants, seeds, invertebrates, amphibians, and fishes in shallow water and mudflats. Low flow H&H zonation changes with flooding, and mobile critters migrate with floodwaters to exploit seasonal habitats. UMR hydrology is well understood with regard to navigation and flood control, but less well understood from an ecological perspective. River hydraulics have been used since the mid-19th Century to shape a deep and stable main navigation channel by clearing and constricting the channel. Large scale flood control and floodplain drainage and development started in the late 19th Century. Low flow river stage regulation to maintain a minimum 9-ft navigation channel was achieved by a network of 37 low-head navigation dams. Upland development has change the rate and volume of runoff that reaches the mainstem. Each of these developments has caused significant ecological change, sometimes with very different effects depending on location along the 1,200 miles of river or the lateral channel-floodplain gradient. River ecologists and conservationists have recommended naturalizing the hydrologic regime with regard to five ecologically relevant characteristics: magnitude, frequency, duration, timing, and rate of change. These parameters are described systematically for the UMR because there is wide availability of river discharge and stage data. Long term discharge and stage data spanning pre and post development are available for nine gauges throughout the system. Each navigation reach also has three or more gauges to assess post dam, riverine to lentic hydrologic gradients in impounded reaches. River stage-discharge modeling has been used to estimate floodplain inundation for flood protection for decades, and the ecological significance of flooding has been generally described, but UMR flood inundation mapping has not been evaluated for ecological significance. Recent flood inundation analysis on high resolution topographic data has greatly increased opportunities to expand pilot study analyses. Historic assessments of flood inundation to assess change due to development has also not been completed, but appropriate map and hydrologic data are widely available and demonstrations have been extremely informative. The five ecologically relevant hydrologic characteristics can also be applied to area of inundation when it is clearly mapped over floodplain elevation data. River hydraulic characteristics operate at fine scales and can vary widely with river discharge. Advanced 2-dimensional hydraulic models developed for many river reaches have been used to design navigation and ecosystem restoration features, and only recently to describe freshwater mussel habitat. These models can be used more widely to estimate habitat availability, material transport capacity, and other ecosystem processes in other river reaches. Keywords: hydrology, hydraulics, inundation mapping, ecohydrology, floodplain river